immediately relevant to game theory 是作主语吗 不是名词形式啊 请详细分析下这句语法

这里用的是倒装
主语：the sex ratios
定语： in certain parasitic wasp species that have a large excess of females.
谓语：are Immediately relevant to game theory
没有倒装的语序为：
the sex ratios in certain parasitic wasp species that have a large excess of females are Immediately relevant to game theory .
在某些雄性黄蜂占多数的黄蜂种类中,其性别比列直接涉及到了博弈论.

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这是个倒装句，真正的主语是the sex ratios in certain parasitic wasp species that have a large excess of females，因为主语太长，顾把主语和之后，避免头重脚轻，有失美感。
若不倒置，则是the sex ratios in certain parasitic wasp species that have a la...