What constitutes a major threat to the survival of turtles a

What constitutes a major threat to the survival of turtles according to Elizabeth Griffin?
此句的主语是what吗

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What constitutes a major threat to the survival of turtles according to Elizabeth Griffin?
根据伊丽莎白·格里芬,什么对海龟的继续生存构成了主要威胁?
疑问词 what 就是这句话的主语,谓语动词是 constitute,其后的 a major threat 是宾语.

1年前

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